but could spread out. 但可以蔓延开来

30. Micro plants, like moss, were first to appear.
    苔藓这样的微型植物率先出现
31. With leaves only one cell thick,
    叶子只有一层细胞
32. they couldn't grow upwards,
    它们无法向上生长
33. but could spread out.
    但可以蔓延开来
34. Reproducing faster than lichen,
    它们的繁殖速度比地衣快
35. they soon carpeted the land,
    很快就铺满陆地
36. which they ruled undisturbed for the next 40 million years.
    在之后的4000万年 不受滋扰地统治着陆地

来自《我们星球上的生命 第一季 第三集 Life on Our Planet S01E03 入侵大地》第 33 行