it was much larger reptiles who would change the course of history. 即将改变历史进程的 是更大的爬行动物

111. lizards are tough and resilient,
     坚韧不拔
112. just like their ancient ancestors.
     就像它们古老的祖先一样
113. But back in the Triassic,
     回到三叠纪
114. it was much larger reptiles who would change the course of history.
     即将改变历史进程的 是更大的爬行动物
115. In the four million years since the Permian extinction...
     二叠纪灭绝之后的400万年中
116. ...Lystrosaurus have dominated.
     水龙兽占据主导地位
117. They emerged into an almost empty world
     它们出现在 一个几乎空无一物的世界中

来自《我们星球上的生命 第一季 第四集 Life on Our Planet S01E04 冷血动物》第 114 行