was

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as it was for their prehistoric ancestors.
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as it was for their prehistoric ancestors. 对于其史前祖先也是如此

But by the time they'd conquered the seas, the ocean was home to deadly predators.
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But by the time they'd conquered the seas, the ocean was home to deadly predators. 但当它们征服大海时 海洋已经成为致命捕食者的家园

was when some of them turned their back on the land
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was when some of them turned their back on the land 也许是其中一些动物放弃陆地

This is an echo of what life was like back in the Triassic.
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This is an echo of what life was like back in the Triassic. 这是三叠纪时期生活的历史回声

...was the age of reptiles.
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...was the age of reptiles. 属于爬行动物

Lystrosaurus was the unwitting dodo of its age.
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Lystrosaurus was the unwitting dodo of its age. 水龙兽成了那个时代的渡渡鸟

it was much larger reptiles who would change the course of history.
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it was much larger reptiles who would change the course of history. 即将改变历史进程的 是更大的爬行动物

another group of survivors was on the rise.
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another group of survivors was on the rise. 又有一群幸存者在崛起

was on the ascent
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was on the ascent 正在崛起

was used to burrowing underground
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was used to burrowing underground 习惯于在地下挖洞